Posts Tagged ‘Erasmus Darwin’

Vestiges: Evidence for Evolution? Part II

 

Charles Darwin attempted to avoid the use of the term “vestiges” largely because the term had been associated with the “erroneous” Larmarckian concept of “use and disuse” that was only “veritable rubbish.”

Jean-Baptiste Lamarck (1744 – 1829) was a member of the French Academy of Sciences and was appointed to the Chair of Botany in 1788. When the Muséum national d’Histoire naturelle was founded in 1793, Lamarck was appointed professor of zoology. In 1801, he published Système des animaux sans vertèbres, a major work on the classifications and coined the term invertebrates. Lamarck is thought to be the first use the term biology in its modern sense. Lamarck continued his work as a premier authority on invertebrate zoology.

Darwin did credit “Lamarck as the first man whose conclusions on the subject excited much attention.… In these works he up holds the doctrine that all species, including man, are descended from other species.”

Lamarck’s theory of evolution, which he referred to as “transformism,” was based on the idea that individuals develop new traits during their own lifetimes by “use and disuse” and transmit them to the next generation. Larmack writes – “Progress in complexity of organization exhibits anomalies here and there in the general series of animals, due to the influence of environment and of acquired habits.”

The giraffe served as Lamarck’s classic example of evolution through “use,” acquiring longer necks in successive generations in competition to reach the ever-scarcer leaves higher in the trees. In illustrating Lamarck’s views on adaptation, Darwin wrote, “To this latter agency he seems to attribute all the beautiful adaptations in nature; such as the long neck of the giraffe for browsing on the branches of trees.”

For Darwin, however, this explanation was simply not scientific – “Lamarck, who believed in an innate and inevitable tendency towards perfection in all organic beings, seems to have felt this difficulty so strongly that he was led to suppose that new and simple forms are continually being produced by spontaneous generation. Science has not as yet proved the truth of this belief.”

One of the most eminent pre-Darwinists was Charles Darwin’s own grandfather, Erasmus Darwin (1731–1802). Erasmus discussed his ideas at length in a two-volume work, Zoonomia, published in 1794. Erasmus wrote that “all … have risen from one living filament.”

Erasmus’ book was widely popular in Western Europe- even translated into German, French, and Italian. Erasmus envisioned that the driving force behind species modification was a result of “lust, hunger, and danger.” In line with Greek philosophy, Erasmus envisioned changes by “continuing to improve its own inherent activity.”

Actually how these “improvements” developed was completely unknown to Lamarck and Erasmus—evolution was a philosophy, not a science. The unknown cause of “improvements” is what drove Darwin to discover the underlying laws of nature—scientifically. Writing in the preface of The Origin of Species, Darwin suggests how Erasmus’s work, although “erroneous,” may have influenced Lamarck: “It is curious how largely my grandfather, Dr. Erasmus Darwin, anticipated the views and erroneous grounds of opinion of Lamarck in his Zoonomia.”

For Lamarck, new characteristics are acquired through the process of “use and disuse.” Darwin’s grandfather, Erasmus Darwin, was a Lamarckian evolutionist. Charles Darwin, however, in pursuit of a “scientific theory” of evolution, initially opposed Lamarckian evolution, only granting the theory marginal support.

In a letter written to J. D. Hooker in 1844, Darwin wrote, “Heaven forefend me from Lamarck nonsense of a ‘tendency to progression.’ … But the conclusions I am led to are not widely diff erent from his, though the means of change are wholly so.” “With respect to books on this subject,” Darwin continues, “I do not know any systematic ones, except Lamarck’s, which is veritable rubbish.”

Although attempting to distance himself from Lamarck’s concepts of “use and disuse” and “vestages,” Darwin distain for “use and disuse” eventually waned as causes for the origin of variation required for the actions of natural selection remained Darwin’s largest unsumountable enigma.

Since then, the term, “vestiges” has once again gained prominence over “rudiments,” as has Larmarckian concepts of evolution. The question remains, however, are structures classified as “vestiges” evidence of evolution? Specifically, have vestiges seemingly lost all or most of their original function in a species through evolution?

To address answers to these questions, we will be examining the most popular example of vestiges—the mammalian appendix in the up-coming posts.   

pre-Origin Notoriety

Charles Darwin recorded in his autobiography that The Origin of Species “is no doubt the chief work of my life. It was from the first highly successful. The first small edition of 1,250 copies was sold on the day of publication, and a second edition of 3,000 copies soon afterwards. Sixteen thousand copies have now (1876) been sold in England and considering how stiff a book it is, this is a large sale.”

While it is unknown how the 1,250 copies could have been sold on “the day of publication” without Amazon.com, what is known is that Darwin was famous long before the publication of the first edition of The Origin of Species in 1859.

Charles Darwin was following in the tradition of his grandfather, Erasmus Darwin—author of the infamous Zoönomia. King George III even asked Erasmus to be his doctor, but he refused the appointment—too busy.

Erasmus was building a vast network of associates that became known as the leading social and philosophical lights. With contacts like Matthew Boulton, Josiah Wedgwood, and James Watt, Erasmus established the Lunar Society that became the main intellectual powerhouse of the Industrial Revolution in England.

By the time Charles Darwin entered Edinburgh University, Zoönomia (meaning “the law of life” in Latin) had become a popular poetry and science textbook. At Edinburgh University, Charles Darwin learned that his professor, Robert Edmund Grant, quoted from Zoönomia for his doctoral thesis. 

Just months after returning from the voyage on the HMS Beagle in February 1837, and before starting working on what is now known as The Origin of Species, Darwin was elected to the Council of the Royal Geographical Society, later accepting Darwin accepted the position of Secretary of the Society in March 1838. Darwin was elected as a Fellow of the Royal Society in January 1839. The Geographic and Royal Society institutions were reserved for the intellectual elite—only.

The Darwin’s in the eighteenth century has been likened to the Kennedy’s of the nineteenth century. Darwin’s notoriety can even be seen at play during the voyage of the HMS Beagle. By British custom, the ship’s surgeon traditionally took the position of the official “naturalist.” Darwin’s role was to be a “gentleman’s naturalist” and assist the ship’s surgeon, Robert McKormick, and Captain FitzRoy. 

On shore in Brazil, however, it was the 22-year old Charles Darwin, not Doctor McKormick, who began receiving all the notoriety and the invitations from dignitaries on shore. Reasonably, McKormick felt upstaged by Darwin. Being sufficiently disgruntled, McKormick left the Beagle at Rio de Janeiro. McKormick’s status was “invalided out” back to Britain.

In 1859, not only was the topic of evolution was “in the air”, Darwin’s word was like E.F. Hutton speaking. The timing was perfect. Darwin’s pre-Origin notoriety preceded the successful launch of one the most influential and contentious books ever in the history of science.

Darwin Mount

The Mount IICharles Robert Darwin was the son of Robert and Susannah Darwin. Darwin was born at the grand family estate known as  in the beautiful “town of flowers,” Shrewsbury, England.  “The Mount” was built by Darwin’s father in 1797 on two and a half acres, now called the “Darwin Gardens.”

Things were run efficiently and orderly at The Mount. Susannah Darwin maintained a “perennial garden diary” to record the details of flowerings and fruiting in the kitchen garden in their pleasure gardens and glasshouses. Darwin was the fifth of six children: three older sisters Marianne, Caroline, and Susanne; one younger sister, Emily Catherine; and an older brother, Erasmus.

As a young child, Darwin was given the nickname of “Babba,” taken from his middle name Robert. As a young teenager, his brother Erasmus just called him “Bobby.” The Darwin home was loving, caring, and cultured. Susannah Darwin skillfully used family teaching moments. When Darwin brought a flower to her, he remembers her saying, “by looking at the inside of the blossom, the name of the plant [can] be discovered;” a lifetime lesson for a budding naturalist.

Zoönomia

Zoonomia

The publication of The Origin of Species by Charles Darwin in 1859 continued the Darwin legacy. Erasmus Darwin, Darwin’s grandfather,  had published the book entitled Zoönomia, or The Laws of Organic Life earlier in 1794. In Zoönomia, Erasmus entertains the basic tenets of evolution and asks the question: 

“Would it be too bold to imagine that all warm-blooded animals have arisen from one living filament, which the great First Cause endued with animality… possessing the faculty of continuing to improve by its own inherent activity, and of delivering down these improvements by generation to its posterity, world without end?”

 As a physician in Lichfield from 1756 to 1781, Erasmus acquired a reputation for being a great healer. He was so successful that King George III asked him to be his doctor, but Erasmus Darwin refused the appointment. Becoming a noted naturalist, writer, poet, and inventor during his own time, Erasmus’ intellectual curiosity eventually led him to be one of the founding members of the Lunar Society. Members of this society were of influence, largely becoming the engine-driving force of the British Industrial Revolution.

 Darwin’s passion to study of nature came into sharper focus during the second year at Edinburgh University. On campus, Darwin became acquainted with Professor Robert Edmund Grant, a proponent of evolution and student of Erasmus Darwin.

 In his doctoral thesis, Grant quoted from Zoönomia. Evolution even at that time was strongly rooted in academic circles. Grant espoused the Lamarckian theory: evolution through acquired characteristics. In his autobiography, Darwin recalls an early conversion with Grant:

 “He one day, when we were walking together he burst forth in high admiration of Lamarck and his views on evolution. I listened without any effect on my mind. Nevertheless it is probable that the hearing rather early in life such views maintained and praised may have favoured my upholding them under a different form in my Origin of Species.”

 In time, Darwin became one of Grant’s keenest students and assisted him with collecting specimens. Grant introduced Darwin to the academic elite of the day, connections that were to become invaluable for his future.



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